Unlike before if a biome says "NONE" there is really not none since there are various global genus groups that inhabit them. This means a much easier start for macro species to migrate there since planktivores, detitivores, herbivores can start right off the bat. Not to mention macro flora have the nitrocycle there so they can easily grow. In short please help out fill in the "NONE" biomes with macro species. Right now most of the empty biomes are rivers and riparian zones. So please help out and fill those biome in with species.
An easy way to find them is to go to the ecosystem page
and use the search button with the term "NONE" to find them all quickly.